Tuppence Middleton (born 21 February 1987) is an English actress known for her performances in film, television and theatre. In 2010, she was nominated for the London Evening Standard Film Awards for Most Promising Newcomer.

Middleton appeared in various films before making her breakthrough in Morten Tyldum's historical drama The Imitation Game (2014), and subsequently appeared in The Wachowskis' science fiction film Jupiter Ascending (2015), Alfonso Gomez-Rejon's historical drama The Current War (2017), David Fincher's film Mank (2020), and the historical drama films Downton Abbey (2019) and Downton Abbey: A New Era (2022).

She made her first television appearance in Bones (2008) and subsequently appeared as a guest in New Tricks (2010), Friday Night Dinner (2011), and Lewis (2013). She also appeared as Jem the Black Mirror episode "White Bear" (2013), as Miss Havisham in Dickensian (2015–2016), as Russian princess Hélène Kuragina in War & Peace (2016), and as Riley "Blue" Gunnarsdóttir in Sense8 (2015–2018).

Early life
Middleton was born in Bristol on 21 February 1987, the daughter of Tina and Nigel Middleton. She has an older sister named Angel and younger brother named Josh. She was named "Tuppence" after the childhood nickname her grandmother gave to her mother. She was raised in Clevedon, Somerset. She has described her younger self as shy, reclusive, and "geeky" at school, but "quite loud and brash" at home; she found youth theatre an "outlet" in which she could be confident. She attended Bristol Grammar School, where she was involved in school plays such as Guys and Dolls. She also attended Stagecoach, a performing arts school in Portishead. She appeared in local drama productions, including a pantomime with her sister at the Princes Hall in Clevedon. She subsequently studied acting at the Arts Educational School in London, earning an honours degree in acting.